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Evolutions that require defeating Alpha or Omega Aniimo
Celestis, Infergon and Sherro all need a boss beaten before they evolve. Here is which fights count, why a wild Wisptis catch doesn't, and the Perfect-only Infergon rule.
The short version
- Celestis evolves into Stellarys at Aniimo level 18, but only after you have defeated an Alpha Stellarys, and the evolution spends a Night Stone.
- Infergon comes only from evolution. Evolving into it starts a fight with Omega Infergon that you have to win, and the evolution needs a Blaze Stone.
- Only a Jawling or Helmwhelp with a Perfect potential rating can become Infergon. Good or Elite quality is not enough.
- When an evolution asks you to defeat wild specimens, such as wild Wisptis, catching them does not count. They have to be defeated.
- A required Alpha is different: beating it marks the criterion as met even if your capture attempt fails, and capturing it counts too.
Most evolutions in Aniimo ask for a level and a stone. A handful ask for something more: you have to beat a particular wild Aniimo first. Sometimes that means several wild specimens of the same Aniimo. Sometimes it means its Alpha form, or its Omega form.
This page covers which evolutions work that way, what counts as a win for each, and the one rule that catches people out most often: capturing the target is not the same as defeating it.
Which evolutions need a defeat

Four lines are gated this way.
Celestis evolution is the one most players run into. Celestis is the Lumin-stage form of its line, and it evolves into its Nova-stage form, Stellarys, once it reaches Aniimo level 18 and you have defeated an Alpha Stellarys to unlock the evolution. The evolution itself spends a Night Stone. Hitting level 18 on its own does nothing until that Alpha is down.
Sherro and Sheldon both need Omega Sherro defeated before they can evolve.
Infergon is gated behind Omega Infergon, with extra conditions covered further down.
Wisptis into Ignitis is the unsettled one. The requirement is given two ways: defeating three wild Wisptis of the same form in succession while using your Wisptis, or a Burning Soul. Which applies is not settled yet, so if one route is not working for you, try the other.
Why capturing does not count, and when it does
When an evolution asks you to defeat wild specimens of an Aniimo, it means defeat. Catching them does not count toward the requirement. The Wisptis route is the clearest case: if you are meant to beat wild Wisptis and you catch them instead, you have made no progress. Knock them out.
Required Alphas follow a looser rule. Defeating the Alpha marks its evolution criterion as met even if your capture attempt afterwards fails, and capturing it also counts. So with an Alpha Stellarys you can throw Anipods without risking the unlock; win the fight and the criterion is yours either way.
The practical difference: for wild specimen counts, fight to the finish and leave your Anipods alone. For an Alpha, a failed catch costs you nothing on the evolution side.
Wild target such as Wisptis
- Capturing does not count toward the defeat requirement
- The wild Aniimo must be defeated
Required Alpha
- Defeating it marks the evolution criterion as met
- It counts even if the capture fails
- Capturing it also counts
Infergon: the Omega fight and the Perfect potential rule

Infergon cannot be caught. It can only be obtained through evolution, and that evolution is the strictest in this group. Infergon has its own guide on this site; here is what matters for the defeat requirement.
Evolving into Infergon starts a fight with Omega Infergon in its quest arena, and you have to win it. Evolving a Helmut into Infergon at the sword triggers that fight, and evolving a Helmwhelp requires the same Omega win. The evolution also needs a Blaze Stone.
The catch comes before any of that. Infergon can only be evolved from a Jawling or Helmwhelp with a Perfect potential rating. Good quality is not enough. Elite quality is not enough either. If you care about Aniimo perfect stats anywhere, this is the place: a pre-evolution that is anything short of Perfect will never become Infergon, however well you fight.

What Alpha bosses give you besides the unlock
Beating an Alpha boss is worth doing even when it is not gating an evolution you need right now. Defeating Alpha bosses yields loot and unlocks evolutionary paths for specific Aniimo, and certain Alphas drop boss-specific materials used for progression and evolution, evolution stones among them. Wave Stones, Gust Stones and Roar Stones all come from defeating Alpha Aniimo, and the first time you defeat an Alpha boss you get a guaranteed set of evolution stones.
Catching an Alpha boss has its own payoff: it gives you an Aniimo that is already evolved, without you having to evolve it yourself. Respawns and which drops to keep are covered in this site's Alpha boss farming guide.
Where to start
Check whether an evolution names a wild specimen, an Alpha or an Omega before you pick a fight. Against wild specimens, defeat them and don't catch them. Against a required Alpha, either result counts, so catch freely. For Infergon, confirm your Jawling or Helmwhelp is Perfect potential and that you have a Blaze Stone before you trigger the Omega Infergon fight.
Most of this page was re-checked against the released game on 2 October 2026. The rest has not been re-confirmed since release.