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How to Unlock Special and Branching Evolutions
Inferlupa needs six Unbroken Flameruff Spirits and a two-day quest. Here is what each special evolution actually asks for: stones, Alphas, gender, place or a trade.
The short version
- Some evolutions lock the materials until a condition is met, so having the stone or resources is not enough on its own.
- Selecting 'how to get' on an evolution requirement in the menu shows where the material or boss is and which Aniimo drop the material.
- Inferlupa needs The Warrior quest finished and six Unbroken Flameruff Spirits; the quest alone does not evolve it and Flux Crystals cannot substitute.
- Celestis evolves into Stellarys at Aniimo level 18, once an Alpha Stellarys has been defeated, and the evolution spends a Night Stone.
- Certain Aniimo only evolve when traded to another player, and others depend on gender, location or a passive talent.
Most Aniimo evolve the plain way: level up, spend the resources, done. A handful don't. Their evolutions ask for something extra first, and until you meet that condition the materials sit in your bag doing nothing.
The extra condition comes in a few flavours. Some need a particular stone. Some need you to beat an Alpha or an Omega boss. Some are tied to a side quest, a gender, a location or a passive talent, and some only trigger when you trade the Aniimo to another player.
This guide goes through each type with the examples that are known, starting with the one that costs the most time.
How locked evolutions work
Evolving any Aniimo needs specific resources. For the special ones, there is a second layer: you must meet a condition beyond the level requirement before the evolution materials can be spent at all. That condition might be a progression milestone, a boss, or something about the Aniimo itself.
The game tells you more than it first appears to. Select 'how to get' on an evolution requirement or material in the menu and it shows where the material or boss can be found, which Aniimo drop the material, and how to obtain it. Check there before you go hunting.
If you bought the Advanced pack, its companion handbook includes evolution materials for the Aniimo family bundled with it, so that line is partly covered already.
Inferlupa: quest first, then the grind
Inferlupa is the clearest case of a quest evolution, and the most demanding. You need to complete the journey quest The Warrior. The Warrior quest chain is time-gated and takes about two real-world days, so start it early.
The catch is that finishing The Warrior alone does not evolve anything. You also need six Unbroken Flameruff Spirits, and Flux Crystals cannot substitute for them.
The Spirits are farmable, but only from one target. They drop from defeating Flameruff the Mighty, the special elite daytime Gamma-stage Flameruff in the Inferlupa elite battle area. Capturing it does not give you one, and plain Flameruffs without the title never drop them. Even when you do beat Flameruff the Mighty, the Spirit is a low-chance drop that comes in place of an evolution stone. Expect repeat fights.
The Warrior also matters outside this evolution. The Beast Fang Ridge leg of the Fragrant Journey needs the Flareburst Flower, and the Flareburst Flower needs The Warrior completed. One quest unlocks both.
six
Unbroken Flameruff Spirits for Inferlupa
Alpha and Omega boss conditions
Certain evolutions require defeating a specific Alpha variant of that species. Others open up only after you beat particular Omega bosses.
Celestis evolution is the textbook Alpha case. Celestis, the Lumin-stage form of its line, evolves into its Nova-stage form Stellarys once it reaches Aniimo level 18 and you have defeated an Alpha Stellarys to unlock the evolution. The evolution then spends a Night Stone, so have one ready before you take the fight.
Bolty into Blazen works on the same idea with a location attached. One step happens at the Blitz Tree in Blitz Wood, standing beneath or on the tree, which is where the Alpha Blazen is fought.
18
Celestis level for Stellarys
Stone evolutions
Stones are the simplest extra requirement, though some come with conditions of their own.
Wisptis evolves into Ignitis with a Night Stone.
Popota into Piopiota needs a Wave Stone, and that is not the whole requirement. You also have to attract an opposite-gender Susuta to follow you. Susuta spawns at night, which is also when you can meet the nighttime forms of both Susuta and Popota, so plan this one for after dark.
Gender, place, talent and trade evolutions
A few evolutions hinge on something about the Aniimo or where you are standing.
Skippy evolves into Glacy, but only if it is female.
Tuckin's mountain form comes from evolving it in Berylline Vale, at a mud pit you reach by flying to Mudstone Pass. Evolve it elsewhere and you will not get that form.
Drapple is not a quest reward. It comes from an exceptional evolution of Nimi when Nimi learns the Dream Song passive talent.
And for anyone asking whether you can trade Aniimo for evolution: yes. Certain Aniimo need to be traded to another player to trigger specific evolutions, so a friend to swap with is part of finishing those lines.
Where to start
If an evolution won't go through, open 'how to get' on the requirement before anything else. Start The Warrior early because of its two-day gate, keep a Night Stone for Celestis, and line up a trading partner for the lines that need one.
Most of this page was re-checked against the released game on 3 October 2026. The rest has not been re-confirmed since release.
